Indian Bluffs State Preserve, Jones Co.
(by Latin species name)

Explanatory Notes

SPECIES
FAMILY
COMMON NAME
ORIGIN
COMMENTS
PTERIDOPHYTES (FERNS & ALLIES)        
Adiantum pedatum L. Adiantaceae Maidenhair fern native  
Asplenium platyneuron (L.) Oakes ex DC. Aspleniaceae Ebony spleenwort native Sandy open woods and thickets.
Asplenium rhizophyllum L. [=Camptosorus rhizophyllus (L.) Link] Aspleniaceae Walking fern native Very common on rocky bluffs and rock outcroppings.
Athyrium filix-femina (L.) Roth var.angustum (Willd.) Moore Aspleniaceae Northern lady fern native  
Athyrium pycnocarpon (Sprengel) Tidestrom [=Diplazium pycnocarpon (Sprengel) Broun] Aspleniaceae Narrow-leaved spleenwort native Lower slopes of rocky bluffs in rich soil, also along streams. Proposed as a special concern species in 1998.
Athyrium thelypterioides (Michx.) Desv. [=Deparia acrostichoides (Sw.) M. Kato ined.] Aspleniaceae Silvery spleenwort native Rich, wooded slopes.
Botrychium dissectum Sprengel f. obliquum (Muhl.) Fern. Ophioglossaceae Oblique grape fern native Both forms usually found together.
Botrychium dissectum Sprengel f. dissectum Ophioglossaceae Dissected grape fern native Both forms usually found together.
Botrychium virginianum (L.) Sw. Ophioglossaceae Rattlesnake fern native  
Cystopteris bulbifera (L.) Bernh. Aspleniaceae Bulblet bladder fern native Rocky bluffs and outcrops.
Cystopteris protrusa (Weath.) Blasdell Aspleniaceae Creeping fragile fern native  
Dryopteris carthusiana (Vill.) H.P. Fuchs [=D. spinulosa (O.F. Muell.) Watt] Aspleniaceae Spinulose wood fern native Base of bluff with northern orientation, rich woodland.
Dryopteris goldiana (Hooker) Gray Aspleniaceae Goldies fern native  
Equisetum arvense L. Equisetaceae Common horsetail native Stream bank, sandy floodplain opening.
Equisetum hyemale L. var. affine (Engelm.) A.A. Eaton Equisetaceae Scouring-rush native Alluvial woods, riparian.
Matteuccia struthiopteris (L.) Todaro var. pensyvanica (Willd.) Morton Aspleniaceae Ostrich fern native Also growing in opening under electrical transmission lines.
Onoclea sensibilis L. Aspleniaceae Sensitive fern native  
Osmunda claytoniana L. Osmundaceae Interrupted fern native Nice, large stands locally.
Pellaea glabella Mett. ex Kuhn Adiantaceae Smooth cliff-brake native Infrequent, steep rock outcroppings, exposed rock in prarie/savanna area.
Polystichum acrostichoides (Michx.) Schott Aspleniaceae Christmas fern native  
Pteridium aquilinum (L.) Kuhn var. latiusculum (Desv.) Underw. ex Heller Dennstaedtiaceae Bracken fern native  
         
GYMNOSPERMS (CONIFERS)        
Juniperus virginiana L. Cupressaceae Red cedar native  
Taxus canadensis Marsh. Taxaceae American yew native Limited to the rims of the larger bluffs where it is locally common.
         
ANGIOSPERMS: DICOTYLEDONS        
Acalypha rhomboidea Raf. Euphorbiaceae Three-seeded mercury native Riparian, sandbars.
Acalypha virginica L. Euphorbiaceae Three-seeded mercury native Bluff top.
Acer negundo L. Aceraceae Box elder native  
Acer saccharinum L. Aceraceae Silver maple native Major componant of the floodplains.
Acer saccharum Marsh. Aceraceae Sugar maple native Common in rich uplands.
Achillea millefolium L. Asteraceae Yarrow native Sandy openings in woods, and pastured areas.
Actaea pachypoda Ell. Ranunculaceae White baneberry native  
Agastache nepetoides (L.) Kuntze Lamiaceae Yellow giant-hyssop native  
Agastache scrophulariifolia (Willd.) Kuntze Lamiaceae Purple giant-hyssop native  
Agrimonia gryposepala Wallr. Rosaceae Agrimony native Open woods, Shrubby areas.
Alliaria petiolata (Bieb.) Cavara & Grande Brassicaceae Garlic mustard Europe Most common along river, following alluvial soil inland. Although absent from some areas, on a whole very well established throughout Preserve.
Amaranthus rudis Sauer Amaranthaceae Amaranth native Riparian (sandbars).
Amaranthus tuberculatus (Moq.) Sauer Amaranthaceae Water hemp native Riparian.
Ambrosia artemisiifolia L. Asteraceae Common ragweed native  
Ambrosia trifida L. Asteraceae Giant ragweed native  
Amelanchier arborea (Michx. f.) Fern. Rosaceae Serviceberry native  
Amorpha canescens Pursh Fabaceae Leadplant native Relict prairie.
Amphicarpaea bracteata (L.) Fern. Fabaceae Hog peanut native  
Anemone canadensis L. Ranunculaceae Canada anemone native Moist soil.
Anemone quinquefolia L. Ranunculaceae Wood anemone native Moist slopes and floodplains.
Anemone virginiana L. Ranunculaceae Tall anemone native  
Antennaria plantaginifolia (L.) Richardson Asteraceae Ladies' tobacco native Top of bluffs with southern exposure, savanna like areas, and pastured areas.
Apios americana Medicus Fabaceae Ground-nut native  
Apocynum androsaemifolium L. Apocynaceae Spreading dogbane native Woodland openings and prairie/savanna area.
Apocynum cannabinum L. Apocynaceae Indian hemp native  
Aquilegia canadensis L. Ranunculaceae Columbine native Usually on rock outcrops.
Arabis glabra (L.) Bernh. Brassicaceae Tower mustard native Sandy pastured areas.
Arabis laevigata (Muhl. ex Willd.) Poiret Brassicaceae Rock cress native Frequently seen in areas of exposed bedrock, on outcrops of rock and shallow soil.
Arabis shortii (Fern.) Gl. Brassicaceae Rock cress native Frequently seen, rock bluffs and outcrops, alluvial soil.
Aralia racemosa L. Araliaceae Spikenard native Infrequent, sheltered areas of northern oriented bluffs.
Arctium minus Bernh. Asteraceae Burdock Eurasia Open disturbed sandy soil, riparian..
Artemisia ludoviciana Nutt. incl. var. gnaphalodes (Nutt.) T&G Asteraceae White sage native Generally found along river banks, and in clearings under high voltage power lines, the var. gnaphalodes however is confined to a sandy pastured area in the SE section.
Asarum canadense L. Aristolochiaceae Wild ginger native  
Asclepias exaltata L. Asclepiadaceae Poke milkweed native Open upland woods.
Asclepias incarnata L. Asclepiadaceae Swamp milkweed native Rare, sandy alluvium.
Asclepias syriaca L. Asclepiadaceae Common milkweed native Open areas of light disturbance.
Asclepias verticillata L. Asclepiadaceae Whorled milkweed native Open areas, not common.
Aster cordifolius L. [=A.cordifolius L. var. sagittifolius (Wedem. ex Willd.) A.G. Jones] Asteraceae Blue wood aster native Upland woods and bluffs.
Aster ericoides L. Asteraceae Heath aster native Woodland edge, dry soil.
Aster lateriflorus (L.) Britton Asteraceae Side-flowered aster native Woodland edge, dry soil.
Aster oblongifolius Nutt. Asteraceae Aromatic aster native Open woods, bluff with southern orientation.
Aster ontarionis Wieg. Asteraceae Ontario aster native Frequently seen on floodplain.
Aster pilosus Willd. Asteraceae Hairy aster native Sandy opening.
Aster shortii Lindley Asteraceae Short's aster native Bluffs.
Barbarea vulgaris R. Br. Brassicaceae Yellow rocket Eurasia Infrequent.
Betula nigra L. Betulaceae River birch native Riparian.
Bidens cernua L. Asteraceae Bur marigold native Riparian.
Bidens connata Muhl. ex Willd. Asteraceae Beggar-ticks native Sandbars and mudflats.
Bidens frondosa L. Asteraceae Beggar-ticks native Riparian.
Blephilia hirsuta (Pursh) Bentham Lamiaceae Woodmint native Common in moist woodland, usually along drainage areas.
Boehmeria cylindrica (L.) Sw. Urticaceae Bog hemp native  
Cacalia muhlenbergii (Sch.-Bip.) Fern. Asteraceae Great indian plantain native  
Calystegia sepium (L.) R. Br. Convolvulaceae Bindweed native Usually in moist open alluvial soil.
Calystegia spithamaea (L.) Pursh Convolvulaceae Low bindweed native Sandy soil of prairie/savanna area. Very common locally.
Campanula americana L. Campanulaceae Tall bellflower native  
Campanula rotundifolia L. Campanulaceae Harebell native Crevices and tops of steep stone bluffs, also exposed rock in prairie/savanna area.
Cannabis sativa L. Moraceae Hemp, Marijuana Eurasia Uncommon, areas of disturbance.
Cardamine penslyvanica Muhl. ex Willd. Brassicaceae Bitter cress native Riparian.
Carduus nutans L. Asteraceae Musk thistle Europe  
Carpinus caroliniana Walter Betulaceae Blue beech native Infrequent to locally common.
Carya cordiformis (Wang.) K. Koch Juglandaceae Bitternut hickory native  
Carya ovata (P. Miller) K. Koch Juglandaceae Shagbark hickory native  
Castilleja coccinea (L.) Sprengel Scrophulariaceae Indian paintbrush native Sandy soil of prairie/savanna area.
Caulophyllum thalictroides (L.) Michx. Berberidaceae Blue cohosh native Rich woodland, also under electric transmission line.
Ceanothus americanus L. var. pitcheri T. & G. Rhamnaceae New Jersey tea native Prairie/savanna area.
Celtis occidentalis L. Ulmaceae Hackberry native  
Cerastium nutans Raf. Caryophyllaceae Nodding chickweed native Exposed hilltops and alluvial valleys.
Chaerophyllum procumbens (L.) Crantz Apiaceae Chervil native Occasional to abundant especially on alluvial soil.
Chenopodium album L. Chenopodiaceae Lamb's quarters Eurasia Rock bluffs, sandy soil.
Chenopodium hybridum L. Chenopodiaceae Maple-leaved goosefoot native Open areas under electric transmission lines.
Circaea lutetiana L. ssp. canadensis (L.) Ascherson & Magnus Onagraceae Enchanter's nightshade native  
Cirsium altissimum (L.) Sprengel Asteraceae Tall thistle native Open ground under electric transmission lines.
Cirsium arvense (L.) Scop. Asteraceae Canada thistle Eurasia Open ground under electric transmission lines.
Cirsium discolor Muhl. ex Willd. Asteraceae Field thistle native Open ground under electric transmission lines, old field.
Cirsium vulgare (Savi) Tenore Asteraceae Bull thistle Eurasia Open ground under electric transmission lines.
Claytonia virginica L. Portulacaceae Spring Beauty native  
Clematis virginiana L. Ranunculaceae Virgin's bower native  
Conza canadensis (L.) Cronq. Asteraceae Horseweed native  
Cornus alternifolia L. f. Cornaceae Alternate-leaved dogwood native  
Cornus drummondii C.A. Meyer Cornaceae Rough-leaved dogwood native Alluvial ground along stream.
Cornus foemina P. Miller ssp. racemosa (Lam.) J.S. Wilson Cornaceae Gray dogwood native Woodland edges.
Corylus americana Walter Betulaceae Hazelnut native  
Crataegus crus-galli L. Rosaceae Cockspur hawthorn native Alluvial woodland
Crataegus margaretta Ashe Rosaceae Hawthorne native Upland woods.
Crataegus punctata Jacq. Rosaceae Hawthorn native Foodplain, and edges of woods.
Crataegus succulenta Schrader ex Link Rosaceae Hawthorn native Floodplain
Cryptotaenia canadensis (L.) DC. Apiaceae Honewort native  
Cuscuta cuspidata Engelm. Convolvulaceae Dodder native Alluvial woods, growing on Aster ontarionis.
Cuscuta gronovii Willd. Convolvulaceae Common dodder native Alluvial woods, growing on Laportea canadensis.
Cuscuta indecora Choisy Convolvulaceae Dodder native Listed as special concern by Iowa DNR. Sandy soil on high river terrace. Growing on Solidago, Ambrosia trifida, etc.
Daucus carota L. Apiaceae Queen Anne's lace Eurasia Rare.
Dentatia laciniata Muhl. ex Willd. Brassicaceae Toothwort native  
Desmodium canadense (L.) DC. Fabaceae Showy tick-trefoil native  
Desmodium glutinosum (Muhl. ex Willd.) Wood Fabaceae Pointed tick-trefoil native  
Dianthus armeria L. Caryophyllaceae Deptford pink Europe Open areas in sandy soil.
Dicentra canadensis (Goldie) Walp. Papaveraceae Squirrel corn native Northern oriented slopes, under rock outcrops in rich, moist soil.
Dicentra cucullaria (L.) Bernh. Papaveraceae Dutchman's breeches native  
Dirca palustris L. Thymelaeaceae Leatherwood native Small, scattered groves at the base of bluffs
Dodecatheon meadia L. Primulaceae Shooting star native Prairie relict, small population. Also on disturbed eastern oriented bluff.
Echinocystis lobata (Michx.) T. & G. Cucurbitaceae Wild cucumber native Open areas under electric transmission lines and riparian.
Elaeagnus umbellata Thunb. Elaeagnaceae Autumn olive e. Asia Sandy open soil under power lines, bluff tops, woodland edges; infrequent.
Ellisia nyctelea L. Hydrophyllaceae Wild tomato native  
Erechtites hieracifolia (L.) Raf. ex DC. Asteraceae Fireweed native Under power lines.
Erigeron philadelphicus L. Asteraceae Fleabane native Riparian.
Erigeron pulchellus Michx. Asteraceae Robin's plantain native Open, southern-oriented bluff, prairie/savanna area. Also pastured bluff.
Erigeron strigosus Muhl. ex Willd. Asteraceae Daisy fleabane native Openings
Euonymus atropurpureus Jacq. Celastraceae Wahoo native Upland woods and bluffs, likely more widespread than noted.
Eupatorium altissimum L. Asteraceae Tall thoroughwort native Sandy soil in clearing under power lines.
Eupatorium purpureum L. Asteraceae Purple joe-pye-weed native  
Eupatorium rugosum Houtt. Asteraceae White snakeroot native  
Euphorbia corollata Engelm. Euphorbiaceae Flowering spurge native Sandy open ground, Prairie relict.
Euphorbia cyathophora Murray Euphorbiaceae Wild poinsettia native Sandy openings and river banks.
Euphorbia hexagona Nutt. ex Sprengel Euphorbiaceae Six-angled spurge native Sandy river banks.
Euphorbia maculata L. Euphorbiaceae spotted spurge native Sandy soil, river banks.
Fragaria virginiana Duchesne Rosaceae Wild strawberry native Sandy opening in woods and prairie/savanna area.
Fraxinus americana L. Oleaceae White ash native Uplands.
Fraxinus nigra Marsh. Oleaceae Black ash native Base and lower slopes of bluffs.
Fraxinus pennsyvanica Marsh. var. lanceolata (Borkh.) Sarg. Oleaceae Green ash native Floodplain.
Galinsoga quadriradiata Ruiz & Pavon Asteraceae Quickweed C&S. Amer. Extremely disturbed soil.
Galium aparine L. Rubiaceae Cleavers native  
Galium boreale L. Rubiaceae Northern bedstraw native Rocky bluffs.
Galium circaezans Michx. Rubiaceae Wild licorice native Bluff tops, infrequent.
Galium concinnum T. & G. Rubiaceae Shining bedstraw native  
Galium triflorum Michx. Rubiaceae Sweet-scented bedstraw native  
Geranium carolinianum L. Geraniaceae Cranesbill native Sandy pastured open areas.
Geranium maculatum L. Geraniaceae Wild geranium native  
Geum canadense Jacq. Rosaceae White avens native  
Geum vernum (Raf.) T. & G. Rosaceae Spring avens native Locally quite common in pastured floodplain woodland. Currently listed as special concern by Iowa DNR.
Glechoma hederacea L. Lamiaceae Creeping Charlie Eurasia Areas of disturbance, riparian.
Gleditsia triacanthos L. Fabaceae Honey locust native  
Gnaphalium obtusifolium L. Asteraceae Everlasting native Sandy openings.
Hackelia deflexa (Wahl.) Opiz Boraginaceae Stickseed native  
Helenium autumnale L. Asteraceae Sneezeweed native Riparian.
Helianthus giganteus L. Asteraceae Tall sunflower native Woodland edges.
Helianthus grosseserratus Martens Asteraceae Sawtooth sunflower native Sandy, alluvial opening.
Helianthus strumosus L. Asteraceae Pale-leaved sunflower native Woodland edges.
Heliopsis helianthoides (L.) Sweet Asteraceae Ox-eye, false sunflower native Prairie/savanna, openings in woodland.
Hepatica nobilis P. Miller var. acuta (Pursh) Steyerm. Ranunculaceae Hepatica native  
Hesperis matronalis L. Brassicaceae Dame's rocket Eurasia Quite frequent on edges of flood plain and some moist alluvial openings where it has established itself quite nicely..
Heuchera richardsonii R. Br. Saxifragaceae Alumroot native Open, southern-oriented bluff, prairie/savanna area.
Hydrophyllum appendiculatum Michx. Hydrophyllaceae Appendaged waterleaf native Northern oriented slopes, under rock outcrops in rich, moist soil.
Hydrophyllum virginianum L. Hydrophyllaceae Virginia waterleaf native  
Hypericum punctatum Lam. Hypericaceae Spotted St. John's wort native Edges of upland woods.
Hypericum pyramidatum Aiton Hypericaceae Giant St. John's wort native Plants noted in pasture next to woodland, and other open areas.
Impatiens capensis Meerb. Balsaminaceae Jewelweed native Riparian.
Impatiens pallida Nutt. Balsaminaceae Yellow jewel weed native  
Isopyrum biternatum (Raf.) T. & G. Ranunculaceae False rue anemone native Carpeting the lower slopes of the bluffs almost exclusively, down to the floodplains.
Juglans cinerea L. Juglandaceae Butternut native  
Juglans nigra L. Juglandaceae Black walnut native  
Krigia biflora (Walter) Blake Asteraceae False dandelion native Open, southern-oriented bluff, prairie/savanna area.
Lactuca biennis (Moench) Fern. Asteraceae Tall blue lettuce native Rare in this part of state, specimen should have been secured.
Lactuca canadensis L. Asteraceae Tall lettuce native Clearings.
Lactuca floridana (L.) Gaertner Asteraceae Blue lettuce native Woodland edges.
Laportea canadensis (L.) Wedd. Urticaceae Wood nettle native On rich slopes and alluvium, this species is hard to ignore when it contacts bare skin.
Leonurus cardiaca L. Lamiaceae Motherwort Asia Disturbed rocky areas, etal.
Lepidium campestre (L.) R. Br. Brassicaceae Field cress Europe Sandy pastured areas.
Lepidium virginicum L. Brassicaceae Poor-man's pepper native Along river bank. Sandy openings.
Lespedeza capitata Michx. Fabaceae Round-headed bush clover native Sandy pastures and prairie/savanna area.
Lindernia dubia (L.) Pennell Scrophulariaceae False pimpernel native Mud flats and sand bars, riparian.
Lithospermum caroliniense (Walter) MacM. Boraginaceae Hairy puccoon native Sandy, open pastures.
Lobelia inflata L. Campanulaceae Indian tobacco native Open ground under electric transmission lines.
Lobelia siphilitica L. Campanulaceae Great lobelia native Open ground under electric transmission lines.
Lobelia spicata Lam. Campanulaceae Pale-spike lobelia native Sandy pastures and prairie/savanna area.
Lonicera dioca L. var.glabrescens (Rydb.) Butters Caprifoliaceae Wild honeysuckle native  
Lonicera morrowii Gray Caprifoliaceae Morrow's honeysuckle Japan Edge of pastured woodland.
Lonicera prolifera (Kirchner) Rehder Caprifoliaceae Wild honeysuckle native  
Lonicera tatarica L. Caprifoliaceae Tararica honeysuckle Eurasia This introduction is not as rampant here, as it sometimes is elsewhere.
Lotus corniculatus L. Fabaceae Bird's-foot trefoil Europe Disturbed openings, pastures.
Lycopus americanus Muhl. ex Barton Lamiaceae Water horehound native Riparian.
Lycopus virginicus L. Lamiaceae Water horehound native Sandbars.
Lysimachia nummularia L. Primulaceae Moneywort Europe Alluvial woodland.
Melilotus officinalis (L.) Pallas Fabaceae Yellow sweet clover Eurasia Disturbed soil of river bank.
Menispermum canadense L. Menispermaceae Moonseed native  
Mentha arvensis L. Lamiaceae Wild mint native Riparian.
Mimulus ringens L. Scrophulariaceae Monkey flower native Riparian.
Mirabilis nyctaginea (Michx.) MacM. Nyctaginaceae Wild four-o'clock native Open, disturbed sandy areas.
Mitella diphylla L. Saxifragaceae Bishop's cap native Common on the larger rock bluffs
Mollugo verticillata L. Aizoaceae Carpet weed Trop. Amer. Moist, disturbed sand.
Monarda fistulosa L. Lamiaceae Wild bergamot native Sandy opening in woods, pastured areas.
Monarda punctata L. Lamiaceae Spotted horsemint native Sandy open area.
Monotropa uniflora L. Ericaceae Indian pipe native Rich upland woods.
Morus alba L. Moraceae White mulberry e. Asia  
Myosoton aquaticum Moench Caryophyllaceae Giant chickweed Europe  
Oenothera laciniata Hill Onagraceae Cut-leaf evening-primrose native Disturbed sandy soil.
Oenothera villosa Thunb. Onagraceae Gray evening primrose native Riparian.
Onosmodium molle Michx. Boraginaceae False gromwell native Prairie/savanna area, rare.
Osmorhiza claytonii (Michx.) C.B. Clarke Apiaceae Sweet cicely native  
Osmorhiza longistylis (Torrey) DC. Apiaceae Anise root native  
Ostrya virginiana (P. Miller) K. Koch Betulaceae Ironwood native  
Oxalis dillenii L. Oxalidaceae Yellow wood sorrel native Sandy, open soil.
Oxalis violacea L. Oxalidaceae Violet wood sorrel native Southern oriented bluffs.
Panax quinquefolia L. Araliaceae Ginseng native Infrequent, rich soil below northern oriented bluffs.
Parthenocissus quinquefolia (L.) Planchon Vitaceae Virginia creeper native  
Parthenocissus vitacea (Knerr) A.S. Hitchc. Vitaceae Woodbine native  
Phlox divaricata L. Polemoniaceae Sweet William, Blue phlox native  
Phryma leptostachya L. Phrymaceae Lopseed native  
Physalis heterophylla Nees Solanaceae Clammy ground cherry native Sandy, deserted agr. ground.
Physostegia parviflora Nutt. ex Gray Lamiaceae False dragonhead native Riparian.
Phytolacca americana L. Phytolaccaceae Pokeweed native  
Pilea pumila (L.) Gray Urticaceae Clearweed native  
Plantago lanceolata L. Plantaginaceae Buckhorn or English plantain Eurasia Disturbed sandy soil.
Plantago rugelii Dcne. Plantaginaceae Common plantain native  
Platanus occidentalis L. Platanaceae Sycamore native  
Podophyllum peltatum L. Berberidaceae Mayapple native  
Polanisia dodecandra (L.) DC. Capparaceae Clammy-weed native Disturbed sandy soil.
Polemonium reptans L. Polemoniaceae Jacob's ladder native  
Polygala senega L. Polygalaceae Seneca snakeroot native Thin soil on southern oriented bluffs, prairie/savanna area.
Polygonum achoreum Blake Polygonaceae Knotweed native Disturbed sandy soil.
Polygonum erectum L. Polygonaceae Knotweed native Disturbed soil.
Polygonum hydropiper L. Polygonaceae Common smartweed native Woodland.
Polygonum hydropiperoides Michx. Polygonaceae Wild water pepper native  
Polygonum lapathifolium L. Polygonaceae Dock-leaved smartweed native Riparian.
Polygonum persicaria L. Polygonaceae Lady's thumb Europe Along river bank.
Polygonum punctatum Ell. Polygonaceae Water-smartweed native  
Polygonum scandens L. Polygonaceae False buckwheat native Woodland edges.
Polygonum virginianum L. Polygonaceae Jumpseed native  
Populus deltoides Bartram ex Marsh. Salicaceae Cottonwood native Alluvial floodplain.
Populus grandidentata Michx. Salicaceae Big-tooth aspen native Wooded slope by prairie/savanna.
Populus tremuloides Michx. Salicaceae Quaking aspen native Upland woods edge.
Potentilla norvegica L. Rosaceae Strawberry weed Eurasia/
native
Opening under powerlines.
Potentilla recta L. Rosaceae Sulfur cinquefoil Europe Sandy opening, disturbed.
Potentilla simplex Michx. Rosaceae Common cinquefoil native Sandy pastured areas and prairie/savanna area.
Prunella vulgaris L. var. lanceolata (Bartram) Fern. Lamiaceae Self heal native Sandy opening in woods, pastured areas.
Prunus americana Marsh. Rosaceae Wild plum native River banks and edges of woodland.
Prunus pensyvanica L. f. Rosaceae Pin-cherry native Rocky bluffs.
Prunus serotina Ehrh. Rosaceae Wild black cherry native  
Prunus virginiana L. Rosaceae Choke cherry native  
Pycnanthemum virginianum (L.) Dur. & Jackson Lamiaceae Mountain mint native Woodland openings.
Pyrola elliptica Nutt. Ericaceae Shinleaf native North orientation on wooded ravine by prairie/savanna.
Quercus alba L. Fagaceae White oak native  
Quercus bicolor Willd. Fagaceae Swamp white oak native Floodplain.
Quercus borealis Michx. f. var. maxima (Marsh.) Ashe [=Quercus rubra L.] Fagaceae Northern red oak native  
Quercus ellipsoidalis E.J. Hill Fagaceae Hill's oak native  
Quercus macrocarpa Michx. Fagaceae Bur oak native Uplands.
Quercus muhlenbergii Engelm. Fagaceae Chinquapin or chestnut oak native On upper portion of bluffs with southern exposure.
Quercus velutina Lam. Fagaceae Black oak native Uplands.
Ranunculus abortivus L. Ranunculaceae Small-flowered crowfoot native  
Ranunculus longirostris Godron Ranunculaceae White water crowfoot native Sandbar and bank of river.
Ranunculus recurvatus Poiret Ranunculaceae Hooked buttercup native Bluff tops, rich woodland.
Ranunculus sceleratus L. Ranunculaceae Cursed crowfoot native Riparian.
Ranunculus septentrionalis Poiret Ranunculaceae Swamp buttercup native  
Ratibida pinnata (Vent.) Barnh. Asteraceae Gray-headed coneflower native Prairie/savanna, Prairie relict.
Rhus glabra L. Anacardiaceae Smooth sumac native Thickets in prairie/savanna area.
Rhus typhina L. Anacardiaceae Staghorn sumac native Sandy clearings along river under power lines.
Ribes cynosbati L. Saxifragaceae Prickly gooseberry native On rock outcrops.
Ribes missouriensis Nutt. ex T. & G. Saxifragaceae Gooseberry native  
Rorippa palustris (L.) Besser Brassicaceae Yellow cress native Intermittent wet areas.
Rosa muliflora Thunb. ex Murray Rosaceae Multiflora rose e. Asia  
Rubus allegheniensis Porter ex Bailey Rosaceae Blackberry native Open ground under electric transmission lines.
Rubus idaeus L. var. strigosus (Michx.) Maxim. Rosaceae Wild red raspberry native Open ground under electric transmission lines.
Rubus occidentalis L. Rosaceae Black raspberry native  
Rubus steelei L.H. Bailey Rosaceae Dewberry native Rocky prairie remnant, prairie/savanna area.
Rudbeckia hirta L. Asteraceae Black-eyed Susan native Pastured openings in woodland, and prairie/savanna area.
Rudbeckia laciniata L. Asteraceae Tall coneflower native Alluvial soil.
Rudbeckia triloba L. Asteraceae Brown-eyed Susan native  
Ruellia humilis Nutt. Acanthaceae Wild petunia native Sandy open areas, not common.
Rumex acetosella L. Polygonaceae Red sorrel Eurasia Sandy opening in woods, sandy pastured areas.
Rumex crispus L. Polygonaceae Curly dock Europe Sandy pastures.
Rumex mexicanus Meisner Polygonaceae Dock native Open area, with prairie grasses under transmission line.
Salix amygdaloides Andersson Salicaceae Peach-leaved willow native  
Salix nigra Marsh. Salicaceae Black willow native Alluvial woodland.
Sambucus canadensis L. Caprifoliaceae Elderberry native  
Sanguinaria canadensis L. Papaveraceae Bloodroot native  
Sanicula canadensis L. Apiaceae Black snakeroot native  
Sanicula gregaria Bickn. Apiaceae Common snakeroot native  
Saponaria officinalis L. Caryophyllaceae Bouncing bet Eurasia cult. Sandy, disturbed soil.
Scrophularia lanceolata Pursh Scrophulariaceae Figwort native Rare along river, and sandy pasture.
Scrophularia marilandica L. Scrophulariaceae Figwort native Especially common in clearing under electrical transmission lines.
Scutellaria lateriflora L. Lamiaceae Mad-dog skullcap native Riparian.
Scutellaria ovata Hill Lamiaceae Skullcap native Woodland.
Senecio pauperculus Michx. Asteraceae Prairie ragwort native Sandy opening in woods, former agricultural use highly probible.
Silene antirrhina L. Caryophyllaceae Sleepy catchfly Europe Sandy old agr. area.
Silene pratensis (Rafn.) Gren. & Godron [=Lychnis alba P. Miller] Caryophyllaceae White campion Europe Sandy pastured areas and openings, rock outcrops, infrequent.
Silphium laciniatum L. Asteraceae Compass plant native Relict prairie.
Silphium perfoliatum L. Asteraceae Cup plant native Alluvial soil
Sisymbrium altissimum L. Brassicaceae Tumble mustard native Disturbed sandy soil, undoubtably more common than noted.
Sisymbrium officinale (L.) Scop. Brassicaceae Hedge mustard Eurasia Opening under electric transmission lines.
Solanum americanum P. Miller Solanaceae Black nightshade native Disturbed soil, riparian..
Solanum carolinense L. Solanaceae Horse nettle native Disturbed sandy areas.
Solidago canadensis L. var. canadensis Asteraceae Tall goldenrod native Openings.
Solidago flexicaulis L. Asteraceae Zig-zag goldenrod native Bluffs and rich woods.
Solidago gigantea Aiton Asteraceae Smooth goldenrod native Woodland edge.
Solidago ulmifolia Muhl. ex Willd. Asteraceae Elm-leaved goldenrod native  
Sonchus oleraceus L. Asteraceae Common sow thistle Europe Opening for electric transmission lines.
Stachys tenuifolia Willd. var. hispida (Pursh) Fern. Lamiaceae Hedge nettle native Riparian.
Staphylea trifolia L. Staphyleaceae Bladdernut native Locally common, forming thickets on rocky slopes and bluffs.
Stellaria graminea L. Caryophyllaceae Common stitchwort Europe Quite common in floodplain pastures with no canopy, less common in woodland edge.
Stellaria media (L.) Vill. Caryophyllaceae Common chickweed native Areas disturbed by moving water
Taraxacum officinale Weber Asteraceae Dandelion Eurasia  
Teucrium canadense L. Lamiaceae Germander native Woodland openings.
Thalictrum dasycarpum Fischer & Ave-Lall. Ranunculaceae Purple meadow-rue native Moist, woodland opening.
Thalictrum thalictroides (L.) Eames & Boivin Ranunculaceae Rue anemone native  
Thaspium barbinode (Michx.) Nutt. Apiaceae Meadow parsnip native Somewhat riparian.
Thlaspi arvense L. Brassicaceae Field penny cress Europe Rare on southern oriented bluff with disturbance.
Tilia americana L. Tiliaceae Basswood native  
Toxicodendron radicans (L.) Kuntze Anacardiaceae Poison ivy native It is possible that T. rydbergii also occurs at this site, but the species were not segregated.
Triodanis perfoliata (L.) Nieuw. Campanulaceae Venus' looking-glass native Open woods and clearings.
Triosteum aurantiacum Bickn. Caprifoliaceae Horse gentian native Open, southern-oriented bluff, and open woods.
Ulmus americana L. Ulmaceae American elm native  
Ulmus rubra Muhl. Ulmaceae Slippery elm native  
Urtica dioica L. Urticaceae Stinging nettle native  
Verbascum thapsus L. Scrophulariaceae Common mullein Europe Rare on rock outcrop, and old field.
Verbena hastata L. Verbenaceae Blue Verbain native Alluvium.
Verbena stricta Vent. Verbenaceae Hoary Vervain native Sandy open soil.
Verbena urticifolia L. Verbenaceae White vervain native Sandy, woodland openings.
Veronia fasciculata Michx. Asteraceae Smooth ironweed native Sandy pasture.
Veronica peregrina L. Scrophulariaceae Speedwell native Sandy soil along river.
Veronicastrum virginicum (L.) Farw. Scrophulariaceae Culver's root native Open woods.
Viola pratincola Greene Violaceae Common blue violet native Woodland openings.
Viola pubescens Aiton Violaceae Yellow violet native  
Viola sororia Willd. Violaceae Hairy blue violet native  
Vitus riparia Michx. Vitaceae Riverbank grape native  
Zanthoxylum americanum P. Miller Rutaceae Prickly ash native Forming small thickets in disturbed woods, both upland and floodplain.
         
ANGIOSPERMS: MONOCOTYLEDONS        
Agropyron repens (L.) Beauv. Poaceae Quackgrass Europe,
Asia
Sandy deserted agr. field.
Agrostis gigantea Roth Poaceae Redtop Europe Pastured areas.
Agrostis perennans (Walter) Tuckerman Poaceae Upland bent native Sandy pasture.
Allium burdickii (Hanes) A.G. Jones [=A. tricoccum Aiton var. burdickii Hanes] Liliaceae Wild leek native Open upland woods, usually noted at top of bluffs.
Allium canadense L. Liliaceae Wild onion naive  
Andropogon gerardii Vitman Poaceae Big bluestem native Fence rows, road ditches.
Arisaema dracontium (L.) Schott Araceae Green dragon native  
Arisaema triphyllum (L.) Schott Araceae Jack-in-the-pulpit native  
Brachyelytrum erectum (Schreber) Beauv. Poaceae Bearded Shorthusk native Open woods on bluff.
Bromus commutatus Schrader Poaceae Hairy chess Eurasia Sandy openings on alluvial plains.
Bromus inermis Leysser Poaceae Smooth brome Europe Disturbed soil by river.
Bromus latiglumis (Shear) A.S. Hitchcock Poaceae Eared brome native  
Bromus pubescens Muhl. ex Willd. Poaceae Canada brome native Rocky wooded areas.
Carex aggregata Mack. Cyperaceae Sedge native Sandy alluvial soil. Currently listed as Special Concern by the Iowa DNR.
Carex albursina Sheldon Cyperaceae White bear sedge native Usually on rich, northern oriented slopes.
Carex amphibola Steudel var. turgida Fern. [=Carex grisea Wahl.] Cyperaceae Sedge native Alluvial soil, infrequent; crevices of rich soil in rocky bluffs.
Carex blanda Dewey Cyperaceae Sedge native  
Carex careyana Dewey Cyperaceae Carey's sedge native Not common, rich wooded slopes. Currently listed as Special Concern by the Iowa DNR.
Carex cephalophora Willd. Cyperaceae Capitate sedge native Openings, open upland woods.
Carex conjuncta Boott Cyperaceae Soft fox sedge native Open alluvial plain, not common.
Carex davisii Schwein. & Torrey Cyperaceae Davis's sedge native  
Carex eburnea Boott Cyperaceae Sedge native Thin soil on crest of rocky bluffs.
Carex gravida Bailey var. lunelliana Mack. Cyperaceae Sedge native  
Carex grayi Carey Cyperaceae Sedge native Rare; alluvial woods.
Carex haydenii Dewey Cyperaceae Hayden's sedge native Colony along edge of river.
Carex hirtifolia Mack. Cyperaceae Hairy sedge native  
Carex hitchcockiana Dewey Cyperaceae Sedge native Open ground under electric transmission lines.
Carex jamesii Schwein. Cyperaceae Grass sedge native Not uncommon.
Carex molesta Mack. Cyperaceae Sedge native Sandy open area on alluvial plain.
Carex muhlenbergii Schkuhr ex Willd. var. muhlenbergii Cyperaceae Muhlenberg's sedge native Sandy open pastured areas.
Carex normalis Mack. Cyperaceae Sedge native Floodplain, broken canopy.
Carex oligocarpa Willd. Cyperaceae Sparse-fruited sedge native Common woodland species.
Carex pedunculata Muhl. ex Willd Cyperaceae Sedge native North oriented slopes, local, but forming large colonies which can account for half the plant cover in a given area.
Carex pensylvanica Lam. Cyperaceae Pennsylvania or oak sedge native Top of southern oriented bluff, under broken canopy. Open woodland.
Carex retroflexa Muhl. ex Willd. Cyperaceae Sedge native Quite common in sandy open areas such as pastures and prairie/savanna area. Possibly, this is only the second documentation of this species in Iowa, the other being in Johnson Co. This plant may be reaching its northern limits here. Special Concern.
Carex rosea Schkuhr. ex. Willd. [=Carex convoluta Mack.] Cyperaceae Sedge native The taxonomy of this group is confusing. This should be the right treatment.
Carex siccata Dewey [=Carex foenea Willd.] Cyperaceae Sedge native This is possibly only the second documentation of this species in Iowa, previously known from a prairie in Clay County in 1936. At this site it occurs in a sandy opening in the woods, and in the prairie/savanna area. Special Concern by the Iowa DNR.
Carex sparganioides Muhl. ex Willd. Cyperaceae Sedge native  
Carex sprengelii Dewey ex Sprengel Cyperaceae Long-beaked sedge native Infrequent on rich slopes, to abundant on sandy river terrace with canopy.
Carex vulpinoidea Michx. Cyperaceae Sedge native Sandy opening.
Cenchrus longispinus (Hackel) Fern. Poaceae Sandbur native Sandy, disturbed soil, including sandbars along river.
Cinna arundinacea L. Poaceae Wood reed native  
Commelina communis L. Commelinaceae Common dayflower e. Asia Sandy river banks.
Corallorhiza odontorhiza (Willd.) Nutt. Orchidaceae Fall coral-root orchid native Upland woods with Triphora trianthophora.
Cyperus aristatus Rottb. Cyperaceae Awned Flat Sedge native Riparian, on sandbars.
Cyperus filiculmis Vahl. Cyperaceae Flat Sedge native Sandy opening.
Cyperus schweinitzii Torrey Cyperaceae Sand Flat Sedge native Sandy open soil under power lines.
Cyperus strigosus L. Cyperaceae Straw-color Flat Sedge native Sandy soil.
Dactylis glomerata L. Poaceae Orchard grass Europe Disturbed areas.
Danthonia spicata (L.) Beauv. ex R. & S. Poaceae Poverty oat grass native Prairie/savanna area.
Diarrhena americana Beauv. var. obovata Gl. Poaceae American beakgrain native Quite common in alluvial woods, occasionally in almost solid stands covering relatively large areas, also common in association with Carex sprengelii.
Dichanthelium acuminatum (Sw.) Gould & Clark var. acuminatum Poaceae Panic grass native Deserted sandy field, and top of southern oriented bluffs.
Dichanthelium acuminatum (Sw.) Gould & Clark var. implicatum (Scribner) Gould & Clark Poaceae Panic grass native Prairie/savanna area, transmission lines, etc.
Dichanthelium latifolium (L.) Gould & Clark Poaceae Broad-leaved panic grass native Open woods and edges.
Dichanthelium oligosanthes (Schultes) Gould var. scribnerianum (Nash.) Gould Poaceae Scribner's panic grass native Open, sandy area under power lines.
Digitaria sanguinalis (L.) Scop. Poaceae Common crabgrass Europe Disturbed soil.
Dioscorea villosa L. Dioscoreaceae Wild yam native  
Echinochloa crusgalli (L.) Beauv. Poaceae Barnyard grass Eurasia Disturbed river banks.
Echinochloa muricata (Beauv.) Fern. var. muricata Poaceae Barnyard grass native Riparian.
Eleusine indica (L.) Gaertner Poaceae Goose grass Old world: Africa? Disturbed sandy soil.
Elymus canadensis L. Poaceae Canada wildrye native High line opening, not common.
Elymus riparius Wieg. Poaceae River bank wild rye native Riparian, growing with Elymus wiegandii.
Elymus villosus Muhl. ex Willd. Poaceae Silky wild rye native  
Elymus virginicus L. Poaceae Virginia wild rye native  
Elymus wiegandii Fern. Poaceae Broad-leaved wild rye native Riparian, growing with Elymus riparius. Also growing on top of of large dislodged bedrock, in pockets of rich soil; steep valley.
Eragrostis frankii C.A. Meyer var. brevipes Fassett Poaceae Sandbar love grass native Sand bars and mud flats.
Eragrostis hypernoides (Lam.) BSP. Poaceae Creeping love grass native Sand bars and mud flats
Eragrostis pectinacea (Michx.) Nees Poaceae Carolina love grass native Disturbed sandy soil, riparian areas.
Eragrostis spectabilis (Pursh) Steudel Poaceae Purple love grass native Dry, open soil.
Erythronium albidum Nutt. Liliaceae Trout-lily, dogtooth-violet native  
Festuca obtusa Biehler Poaceae Nodding fescue native  
Galearis spectabilis (L.) Raf. Orchidaceae Showy orchid native  
Hypoxis hirsuta (L.) Cov. Liliaceae Yellow stargrass native Top of southern oriented bluffs.
Hystrix patula Moench [=Elymus hystrix L.] Poaceae Bottle-brush grass native  
Juncus tenuis Willd. Juncaceae Pathrush native Sandy openings.
Leersia virginica Willd. Poaceae White grass native  
Luzula multiflora (Retz.) Lej. Juncaceae Woodrush native Open woodland on southern oriented bluff, growing around trees in open soil w/ Carex pensylvanica
Melica nitens (Scribner) Nutt. ex Piper Poaceae Three-flowered melic grass native Sandy openings in alluvial woods.
Milium effusum L. Poaceae Millet-grass native Apparently documented only once in Iowa, from Allamakee county. At this preserve the grass was collected in an artificial clearing that is kept free of trees for electric power lines. Currently listed as Special Concern by the Iowa DNR.
Muhlenbergia frondosa (Poiret) Fern. Poaceae Wirestem muhly native Riparian
Muhlenbergia mexicana (L.) Trin. Poaceae Wirestem muhly native Tops of southern oriented bluffs.
Muhlenbergia schreberi J.F. Gmelin Poaceae Nimblewill native Bluff tops, and dry open soil.
Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. Poaceae Rock muhly native Southern oriented bluff tops.
Muhlenbergia sylvatica (Torrey) Torrey ex Gray Poaceae Forest muhly native Open woods.
Oryzopsis racemosa (Smith) Ricker Poaceae Black seeded rice-grass natve Rich soiled slopes and bluffs of any orientation.
Panicum capillare L. Poaceae Common witchgrass native Disturbed soil, sometimes riparian.
Panicum dichotomiflorum Michx. Poaceae Fall panic grass native Sandy river banks.
Panicum virgatum L. Poaceae Switchgrass native Sandy openings under electrical transmission lines.
Paspalum setaceum Michx. var. muhlenbergii (Nash) D. Banks Poaceae Bead-grass native Sandy open soil.
Phalaris arundinacea L. Poaceae Reed canary grass native Sometimes quite abundant on floodplain.
Phleum pratense L. Poaceae Timothy Europe Disturbed areas, pastures.
Poa pratensis L. Poaceae Kentucky bluegrass Europe Sandy opening in woods, and pastured areas.
Poa sylvestris Gray Poaceae Woodland bluegrass native Rich wooded slopes.
Polygonatum biflorum (Walter) Ell. Liliaceae Solomon's seal native  
Potamogeton nodosus Poiret Potamogetonaceae Pondweed native Farm impoundment.
Schizachne purpurascens (Torrey) Swallen Poaceae False medic native Not common; in pockets of soil on rock outcrops.
Setaria glauca (L.) Beauv. Poaceae Yellow foxtail Europe Disturbed soil, riparian at times.
Sisyrinchium campestre Bickn. Iridaceae Blue-eyed grass native Top of southern oriented bluffs, prairie/savanna area.
Smilacina racemosa (L.) Desf. Liliaceae False Solomon's seal native  
Smilax ecirrhata (Engelm. ex Kunth) S. Watson Liliaceae Carrion flower native  
Smilax herbacea L. Liliaceae Carrion flower native  
Smilax hispida Muhl. Liliaceae Greenbriar native  
Sorghastrum nutans (L.) Nash Poaceae Indian grass native Road ditch.
Sphenopholis obtusata (Michx.) Scribner var. obtusata Poaceae Wedge grass native Prairie/savanna area.
Sphenopholis obtusata (Michx.) Scribner var. major (Torrey) K.S. Erdman [=S. intermedia Rydb.] Poaceae Wedge grass native Open woods on bluff.
Tradescantia ohiensis Raf. Commelinaceae Spiderwort native Sandy, woodland openings.
Trillium flexipes Raf. Liliaceae Nodding trillium native Northern oriented slopes, under rock outcrops in rich, moist soil.
Triphora trianthophora (Sw.) Rydb. Orchidaceae Nodding pogonia native Large population under dense canopy in upland woods, few herbaceous plants, with much leave mulch.
Uvularia grandiflora Small Liliaceae Bellwort native  
Zigadenus glaucus Nutt. Liliaceae White camass native Crevices and tops of steep stone bluffs, northern orientation.

TOTAL: 448 species
Alien species: 12%, but this is not a comprehensive list. Naturalized species in old fields are not included.
       

Last Updated: 04/08/2002

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